EAR. (=E).
Anatomically
E. is composed of 3 parts: 1) Outer E., 2) Middle E., 3) Inner E.
INNER E. contains 3 types of tubular components: 1) Cochlear duct, 2) Semicircular duct, 3) Saccule and Utricle. Each tubule contains proper TISSUE SENSORY ZONE. COCHLEAR duct has 3 walls: a) Lateral wall – is
stria vascularis, b) Medial wall – is
vestibular (or Reissners)
membrane, c) Lower wall – is
basilar membrane. It is place for location of sensory zone, named
organ of
CORTI (or organ of
HEARING).
ORGAN of
CORTI (or organ of
hearing) is composed of:
-
HAIR SENSORY EPITHELIAL cells,
-
Supporting epithelial cells,
-
Inner tunnel,
-
TECTORIAL membrane.
-
Afferent nerve fibres of cochlear nerve.
There are
2 types of
hair sensory cs: A) Outer
hair sensory cs. It are CYLINDRICAL cs with apical stereocilia. B) Inner
hair sensory cs. It are FLASK-shaped cs with apical stereocilia. Nerve fibres of cochlear nerve form
synaptic junction with
basal surface of hair cs.
Hairs’ means location of
STEREOCILIA on apical surface. Stereocilia – are modified
microvilli.
Funct.:
Hair cs (and
organ of
CORTI, as sensory zone) transform energy of sound waves into nerve impulse, providing
sense of
HEARING.
Injury of
HAIR cs in organ of Corti leads to
DEAFNESS.
Supporting epithelial cells in organ of Corti include:
-
Inner and outer phalangeal cs, locating beneath hair cs. It have long phalangeal process.
-
Inner and outer pillar cs. It form wall of inner tunnel. Tunnel contains ENDOLYMPH and afferent nerve fibres of cochlear nerve.
Organ of BALANCE (vestibular apparatus):
It contains
2 tissue
sensory zones:
-
CRISTAE AMPULLARES (of semicircular duct)
-
MACULAE (of saccule and of utricle).
MACULAE are composed of:
-
HAIR SENSORY EPITHELIAL cells,
-
Supporting epithelial cells,
-
OTOLITHIC membrane.
-
Afferent nerve fibres of cochlear nerve.
There are
2 types of
hair sensory cs:
-
Type 1 hair sensory cs. It are FLASK-shaped cs with 2 types of apical SPECIAL organelles: a) short non motile stereocilia and b) single LONG k i n o c i l i u m. Kinocilium – is motile cilium.
-
Type 2 hair sensory cs. It are CYLINDRICAL cs with same 2 apical special organelles.
Stereocilia and kinocilium are
embedded in gelatinous
OTOLITHIC membrane. Membr. is composed of glycoproteins and crystals known as OTOLITHS (calcium carbonate). Funct.:
Hair sensory cs (and
MACULA, as sensory zone) transform energy of
LINEAR movements,
GRAVITY and
VIBRATION into nerve impulse, providing perception of all these irritations.
CRISTA AMPULLARIS is composed of:
-
HAIR SENSORY EPITHELIAL cells,
-
Supporting epithelial cells (WITHOUT differentiation on types),
-
CUPULA gelatinosa,
-
Afferent nerve fibres of cochlear nerve.
There are
2 types of
hair sensory cs in crista amullaris:
-
Type 1 hair sensory cs. and
-
Type 2 hair sensory cs
Both types have
SAME characteristic, as in
macula.
Stereocilia and kinocilium of hair cs are
embedded in CUPULA gelatinosa. Funct.:
Hair sensory cs (and
CRISTA AMPULLARIS , as sensory zone) transform energy of
ANGULAR movements into nerve impulse, providing perception of this irritation.
Organ of
OLFACTION (is located in olfactory epithelium of nasal cavity) is composed of: 1) OLFACTORY RECEPTOR cs (=
ORC), 2) Tall sustentacular cs, 3) Short basal cs
ORC is BIPOLAR
NEURON (compare: in organ of Corti sensory cs – are EPITHELIUM).
Modified dendrite forms dilation, named
olfactory vesicle. 6-8
olfactory cilia extend from olfactory vesicle. Funct.: Olfactory cilia of
ORC provide
perception of
odors. During this ORC generates nerve impulse and conducts it along basal axon into brain.
Basal cs- is origin for
regeneration of ORCs.
Составитель – доцент В.В. Бондаренко.